- Grug as Lightning McQueen
- Belt as Mater
- Eep as Sally
- Finn McMissile (from Cars 2) as Doc Hudson
- Rocky (from Chicken Run) as Sheriff
- Guy as Luigi
- Jethro (from Olly the Little White Van) as Guido
- Donkey (from Shrek) as Ramone
- Ugga as Flo
- Muck (from Bob the Builder) as Fillmore
- Astronaut Al (from Engle Benjy) as Sarge
- Gran as Lizzie
- Cookie (from Atlantis: the Lost Empire) as Red
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